Measurement and modeling of spatially explicit variation in light transmission through interior cedar-hemlock forests of British Columbia

We have characterized canopy geometry and light transmission by the nine dominant conifer and broad-leaved tree species of the interior cedar-hemlock (ICH) forests of northern British Columbia. Our field data were used to parameterize a spatially explicit model of light transmission through mixed-species forests. That model, a component of the forest dynamics simulator SORTIE, was developed for eastern deciduous forests, and this paper presents a test of that model in a very different ecosystem. Our results show that individual crowns of the ICH forests intercepted much more light than species of eastern deciduous forests but that the canopy as a whole allowed greater light penetration, largely because of openings between the relatively narrow, conical crowns of the western conifers. Light transmission by individual crowns was correlated with shade tolerance among the conifers (as in eastern deciduous species), but crown depth was not (in contrast with eastern species). Despite the fundamental differences in the nature of light transmission in the two ecosystems, the SORTIE light model developed for eastern deciduous forests was effective at predicting spatial variation in understory light levels in these western coniferous forests. The goodness of fit of such a simple model suggests that the most important factors regulating spatial variation in understory light levels in these forests are simply the sizes and distribution of nearby trees, and the local sky brightness distribution. Discrete canopy gaps represent a special case in which a region of the canopy is not occupied by crowns.