[1] Univ Tennessee, Hlth Sci Ctr, Dept Neurol, Memphis, TN 38163 USA
[2] Vet Affairs Med Ctr, Res Serv, Memphis, TN USA
[3] Univ Tennessee, Hlth Sci Ctr, Ctr Neurobiol Brain Dis, Memphis, TN USA
[4] Univ Tennessee, Hlth Sci Ctr, Dept Anat & Neurobiol, Memphis, TN USA
[5] Univ Tennessee, Hlth Sci Ctr, Ctr Excellence Connect Tissue Dis, Memphis, TN USA
[6] Univ Washington, Dept Neurol, Seattle, WA USA