[1] Rural Dev Adm, Dept Agr Biotechnol, Natl Acad Agr Sci, Suwon 441707, South Korea
[2] Seoul Natl Univ, Dept Plant Sci, Coll Agr & Life Sci, Seoul 151921, South Korea
[3] Seoul Natl Univ, Natl Instrumentat Ctr Environm Management, Coll Agr & Life Sci, Seoul 151921, South Korea
[4] Rural Dev Adm, Vegetable Res Div, Natl Inst Hort & Herbal Sci, Suwon 441440, South Korea
[5] Korea Res Inst Biosci & Biotechnol, Taejon 305806, South Korea
[6] Kyungpook Natl Univ, Sch Appl Biosci, Coll Agr & Life Sci, Taegu 702701, South Korea
[7] Chungnam Natl Univ, Dept Hort, Taejon 305764, South Korea
[8] Norwich Res Ctr, John Innes Ctr, Norwich NR4 7UH, Norfolk, England