[1] Univ Massachusetts, Dept Biochem & Mol Biol, LGRT, Amherst, MA 01003 USA
[2] Univ Massachusetts, Cell & Mol Biol Program, LGRT, Amherst, MA 01003 USA
[3] Univ Massachusetts, Plant Biol Grad Program, LGRT, Amherst, MA 01003 USA
[4] Univ Leicester, Dept Biol, Leicester LE1 7RA, Leics, England