[1] Japan Agcy Marine Earth Sci & Technol, Inst Res Earth Evolut, Yokosuka, Kanagawa 2370061, Japan
[2] Shizuoka Univ, Sch Sci, Dept Life & Earth Sci, Shizuoka, Japan
[3] U Dom Corp, Tokyo, Japan
[4] Nagasaki Univ, Fac Fisheries, Nagasaki 852, Japan
[5] Southampton Oceanog Ctr, DEEPSEAS Benth Biol Grp, Southampton SO143 ZH, Hants, England