[1] Wageningen Univ Agr, Lab Theoret Prod Ecol, Grp Plant Prod Syst, NL-6700 AK Wageningen, Netherlands
[2] INRA, Unite Agron, F-31326 Castanet Tolosan, France
[3] Wageningen Univ Agr, Sub Dept Math, NL-6703 HA Wageningen, Netherlands
[4] Wageningen Univ Agr, Grp Biol Farming Syst, NL-6709 PG Wageningen, Netherlands