[1] Louisiana State Univ, Coastal Fisheries Inst, Dept Oceanog & Coastal Sci, Baton Rouge, LA 70803 USA
[2] Univ N Carolina, Dept Marine Sci, Chapel Hill, NC 27599 USA
[3] Natl Marine Fisheries Serv, Alaska Fisheries Sci Ctr, Seattle, WA 98115 USA
[4] Japan Agcy Marine Earth Sci & Technol, Frontier Res Ctr Global Change, Ecosyst Change Res Program, Kanazawa Ku, Kanagawa 2360001, Japan
[5] Hokkaido Univ, Frontier Res Ctr Global Change, Ecosyst Change Res Program, Kita Ku, Sapporo, Hokkaido 0600810, Japan
[6] Hokkaido Univ, Grad Sch Environm Earth Sci, Kita Ku, Sapporo, Hokkaido 0600810, Japan
[7] Fisheries & Oceans Canada, Pacific Biol Stn, Nanaimo, BC V9R 5K6, Canada
[8] Alaska Dept Fish & Game, Div Commerical Fisheries, Kodiak, AK 99615 USA