[1] INRA, UHP Interact Arbres Microorganismes, Unite Mixte Rech, F-54280 Seichamps, France
[2] Univ Kitakyushu, Kitakyushu, Fukuoka 8080135, Japan
[3] Univ Freiburg, Inst Biol 2, D-79104 Freiburg, Germany
[4] Univ Paris 07, LEM, EA 3514, F-75251 Paris 05, France