[1] Gunma Univ, Inst Mol & Cellular Regulat, Cell Biol Lab, Gunma 3718510, Japan
[2] Electrotech Lab, Supermol Sci Div, Tsukuba, Ibaraki 3058568, Japan
[3] Nagoya Univ, Sch Med, Dept Physiol, Aichi 4668550, Japan
[4] Tokyo Gakugei Univ, Dept Biol, Tokyo 1848501, Japan