THE RELATIONSHIP BETWEEN CLIMATIC VARIABLES AND WOOD STRUCTURE IN PINUS-HALEPENSIS MILL .1. EFFECTS OF HUMIDITY AND TEMPERATURE

The weekly increments under various conditions of a dozen of 30 to 60 years old Halep pines grown in the Lecce area have been measured during two years with a band dendrometer. The data have been compared with the X-ray density of a stem in radial direction, the weekly precipitations and the integrated temperature excess above 8°C. We have found that growth rate behaviour is practically the same for all the pines considered. This indicate a strict dependence of growth on external factors such as climate and other environmental conditions. Vegetation activity does not seem to have regular dormancy periods: it stops when temperature and humidity become prohibitive and resumes as soon as they become favourable. The volume of wood formed in late-season (July-January) equals more or less that of early-season (February-June), whereas the wood mass and the number of cells in late-season is nearly double of that in early-season. During periods of high cambial activity one observes a lower cell lignification. However, on the whole, the photoperiodicity hypothesis in lignification seems to be confirmed. The growth rate appears inversely related to density. The lignification process appears to extend to the outer xylem layer of 1-2 mm and seems most effective during dry and warm seasons when cambial activity is low. Density wriggles appear to derive from combined variations of cambial activity, cell size and lignification, mostly corresponding to environmental events such as periods of drought or of cold weather. Finally, during the dry seasons we find that a precipitation event is generally followed within a time less than a week, by an increase of growth rate. Whereas during the wet season the growth rate variation is very similar to that of temperature excess. © 1990 Springer-Verlag.