The chronological sequence of Middle and Late Sarmatian fossil mammalian localities of Ukraina, Moldova, and Transcaucasus is presented. The stratigraphic position of the localities is indicated within the substages of the Eastern Paratethys and within the magnetochronological scale. The localities of the Bessarabian substage date to the second half of the polarity Chron 11 and first half of Chron 10. The fauna of these localities corresponds to that of the Early Vallesian (MN9). The localities of the Khersonian substage date to the second half of Chron 9 (Anomaly 5) and the beginning of Chron 8. The fauna is correlated with that of the Late Vallesian (MN10). The lower boundary of the Vallesian is close to the base of the Bessarabian. The MN9/MN10 boundary is not older than the boundary between the Bessarabian and the Khersonian substages. The upper boundary of the Vallesian nearly coincides with the boundary between the Sarmatian and the Meotian.