TREE POPULATION-DYNAMICS, GROWTH, AND MORTALITY IN OLD-GROWTH FORESTS IN THE WESTERN OLYMPIC MOUNTAINS, WASHINGTON

Tree population dynamics, growth, and mortality were determined in old-growth forested watersheds in the Hoh River valley, Olympic Peninsula, Washington: West Twin Creek (elevation 180-850 m) and Hoh Lake (elevation 1250-1525 m). Principal tree species at West Twin Creek are Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), western hemlock (Tsuga heterophylla (Raf.) Sarg.), Pacific silver fir (Abies amabilis (Dougl.) Forb.), western red cedar (Thuja plicata D. Don), and Sitka spruce (Picea sitchensis (Bong.) Carr.). At Hoh Lake they are Pacific silver fir, mountain hemlock (Tsuga mertensiana (Bong.) Carr.), and Alaska cedar (Chamaecyparis nootkatensis (D. Don) Spach). In 1985 stem densities for trees >5 cm DBH averaged 476 and 489 ha-1 in the upper and lower West Twin Creek watershed, respectively, and 508 ha-1 at Hoh Lake watershed. Stem densities at both sites declined 3-5% from 1985 to 1990. Western hemlock and Pacific silver fir were the dominant species at West Twin Creek and Hoh Lake, respectively. From 1985 to 1990 the annual mortality rate was 0.8% at West Twin Creek and 0.9% at Hoh Lake. Pacific silver fir had the highest mortality rate. No Douglas-fir or western red cedar trees died. The primary causes of mortality were as follows: suppression-unknown, diseases, insects, and windthrow. In 1985 basal areas were 77, 87, and 94 m2.ha-1 in the lower and upper West Twin Creek watershed and Hoh Lake watershed, respectively. There was a 5% increase and a 1% decrease in basal area from 1985 to 1990 in the lower and upper West Twin Creek watershed, respectively, and a 4% decrease at Hoh Lake. Western red cedar, Douglas-fir, and western hemlock increased in basal area, while the other species declined.