[1] Concordia Univ, Dept Geog, Montreal, PQ H3G 1M8, Canada
[2] Inst Ecosyst Studies, Millbrook, NY 12545 USA
[3] British Columbia Forest Serv, Res Sect, Smithers, BC V0J 2N0, Canada
[4] Univ Quebec, Grp Rech Ecol Forestiere Interuniv, Montreal, PQ H3C 3P8, Canada