[1] James Cook Univ N Queensland, Dept Zool & Trop Ecol, Townsville, Qld 4811, Australia
[2] Univ Queensland, Trop Hlth Program, Mol Parasitol Unit, Australian Ctr Int & Trop Hlth & Nutr, Brisbane, Qld 4029, Australia
[3] Queensland Inst Med Res, Brisbane, Qld 4029, Australia
[4] Univ Grenoble 1, UMR CNRS 5553, Lab Biol Populat Altitude, F-38041 Grenoble 09, France