[1] Univ Leeds, Ctr Plant Sci, Leeds LS2 9JT, W Yorkshire, England
[2] Univ York, Dept Biol, CNAP, York YO10 5YW, N Yorkshire, England
[3] Univ Nottingham, Div Agr & Environm Sci, Loughborough LE12 5RD, Leics, England
[4] Univ Western Australia, ARC Ctr Excellence Plant Energy Biol, Nedlands, WA 6009, Australia
[5] Rothamsted Res, Crop Performance & Improvement Div, Harpenden AL5 2JQ, Herts, England