[1] Kyoto Univ, Fac Med, Dept Cell Biol, Kyoto 6068501, Japan
[2] Jikei Univ, Sch Med, Inst DNA Med, Dept Mol Cell Biol, Tokyo 1058461, Japan
[3] Osaka Univ, Biol Sci Lab, Grad Sch Frontier Biosci, Suita, Osaka 5650871, Japan
[4] Osaka Univ, Biol Sci Lab, Grad Sch Med, Suita, Osaka 5650871, Japan
[5] Kobe Univ, Grad Sch Med, Dept Physiol & Cell Biol, Div Cell Biol, Kobe, Hyogo 6500017, Japan
[6] Japan Sci & Technol Agcy, PRESTO, Kawaguchi, Saitama 3320012, Japan