[1] NIA, Lab Mol Gerontol, NIH, Baltimore, MD 21224 USA
[2] NICHHD, Lab Genom Integr, NIH, Bethesda, MD 20892 USA
[3] Osaka Univ, Grad Sch Frontier Biosci, Osaka, Japan
[4] Erasmus MC, Ctr Biomed Genet, MGC Dept Cell Biol & Genet, Rotterdam, Netherlands
[5] Japan Sci & Technol Agcy, Solut Oriented Res Sci & Technol, Wako, Saitama, Japan
[6] RIKEN, Discovery Res Inst, Wako, Saitama 35101, Japan