[1] Natl Ctr Neurol & Psychiat, Natl Inst Neurosci, Dept Inherited Metab Dis, Kodaira, Tokyo 1808502, Japan
[2] Nihon Univ, Coll Bioresource Sci, Dept Appl Biol Sci, Fujisawa, Kanagawa 2520813, Japan
[3] Natl Ctr Neurol & Psychiat, Natl Inst Neurosci, Dept Biochem & Cellular Biol, Kodaira, Tokyo 1808502, Japan
[4] RIKEN, Inst Phys & Chem Res, Tsukuba Life Sci Ctr, Mol Neurobiol Lab, Tsukuba, Ibaraki 3050074, Japan
[5] Univ Tokyo, Inst Med Sci, Dept Mol Neurobiol, Tokyo 1080071, Japan