[1] Washington Univ, Sch Med, Dept Med, St Louis, MO 63110 USA
[2] Washington Univ, Sch Med, Dept Mol Microbiol, St Louis, MO 63110 USA
[3] Washington Univ, Sch Med, Dept Pathol & Immunol, St Louis, MO 63110 USA
[4] Yale Univ, Sch Med, Rheumatol Sect, New Haven, CT 06520 USA
[5] Yale Univ, Sch Med, Immunobiol Sect, New Haven, CT 06520 USA