[1] Inst Nat Conservat, B-1070 Brussels, Belgium
[2] Ctr Estuarine & Coastal Ecol, Netherlands Inst Ecol, NL-4400 AC Yerseke, Netherlands
[3] RIKZ, Natl Inst Coastal & Marine Management, NL-4330 EA Middelburg, Netherlands
[4] Univ Antwerp, Dept Biol, B-2610 Antwerp, Belgium