[1] Bournemouth Univ, Ctr Conservat Ecol & Environm Change, Poole BH12 5BB, Dorset, England
[2] Univ Veracruzana, Inst Invest Forestales, Xalapa 91000, Veracruz, Mexico
[3] Univ Concepcion, Fac Ciencias Forestales, Concepcion, Chile
[4] Univ Nacl Comahue, Lab Ecotono, Inst Invest Biodiversidad & Medio Ambiente, Consejo Nacl Invest Cient & Tecn, RA-8400 San Carlos De Bariloche, Rio Negro, Argentina
[5] Univ Concepcion, Dept Econ, Concepcion, Chile