[1] Kobe Univ, Fac Sci, Dept Biol, Kobe, Hyogo 657, Japan
[2] Kobe Univ, Grad Sch Sci & Technol, Kobe, Hyogo 657, Japan
[3] Wayne State Univ, Sch Med, Dept Ophthalmol, Detroit, MI 48201 USA
[4] Wayne State Univ, Sch Med, Dept Pharmacol, Detroit, MI 48201 USA
[5] Wayne State Univ, Sch Med, Kresge Eye Inst, Detroit, MI 48201 USA
[6] Nagoya Univ, Sch Med, Dept Anat, Nagoya, Aichi 466, Japan