[1] Univ Michigan, Sch Med, Div Med & Mol Genet, Ann Arbor, MI 48104 USA
[2] Univ Michigan, Sch Publ Hlth, Ann Arbor, MI 48109 USA
[3] Johns Hopkins Univ, Sch Hyg & Publ Hlth, Baltimore, MD USA
[4] Johns Hopkins Univ, Johns Hopkins Oncol Ctr, Baltimore, MD USA
[5] Howard Hughes Med Inst, Baltimore, MD USA