[1] Queensland Inst Med Res, Conjoint Gastroenterol Lab, Herston, Qld 4029, Australia
[2] Queensland Inst Med Res, Human Genet Lab, Herston, Qld 4029, Australia
[3] Okayama Univ, Grad Sch Med & Dent, Dept Surg Gastroenterol, Okayama 7008530, Japan
[4] Univ Melbourne, Ctr Genet Epidemiol, Parkville, Vic 3052, Australia
[5] McGill Univ, Dept Pathol, Montreal, PQ, Canada