[1] Univ Maryland, Dept Geog, College Pk, MD 20742 USA
[2] Woods Hole Res Ctr, Woods Hole, MA 02543 USA
[3] Univ Oxford, Dept Zool, Trypanosomiasis & Land Use Africa Res Grp, Oxford OX1 3PS, England
[4] Kenya Govt Med Res Ctr, Wellcome Trust Collaborat Programme, Nairobi 00100, GPO, Kenya