[1] Kyushu Univ, Grad Sch Sci, Dept Biol, Higashi Ku, Fukuoka 8128581, Japan
[2] Kyushu Univ, Grad Sch Med Sci, Dept Mol Biol, Higashi Ku, Fukuoka 8128582, Japan
[3] Kyushu Univ, Med Inst Bioregulat, Lab Embryon & Genet Engn, Dept Mol & Cellular Biol,Higashi Ku, Fukuoka 8128582, Japan
[4] Osaka Univ, Inst Prot Res, Div Prot Metab, Suita, Osaka 5650871, Japan