[1] Univ Tokyo, Lab Wildlife Biol, Bunkyo Ku, Tokyo 1138657, Japan
[2] Russian Acad Sci, Inst Biol & Soil Sci, Vladivostok 690022, Russia
[3] WING, Wild Bird Soc Japan, Hino, Tokyo 1910041, Japan
[4] Yamashina Inst Ornithol, Abiko, Chiba 2701145, Japan
[5] Khingansky Nat Reserve, Arkhara, Amursk Region, Russia
[6] NTT, Wireless Syst Labs, Yokosuka, Kanagawa 2380313, Japan