Glypholoma larvae at last:: phylogenetic implications for basal Staphylinidae?: (Coleoptera: Staphylinidae: Glypholomatinae)

Species of Glypholoma Jeannel, a genus previously moved to Staphylinidae from Silphidae, have adult synapomorphies of the Omaliine Group, a probably basal lineage of about 10 000 species. Adult characters have not definitively resolved the position of Glypholomatinae within the Omaliine Group, but suggest that it may be relatively basal. Larvae and pupae recently collected with adults of Glypholoma pustuliferum Jeannel are attributable to that species on the basis of that association, a good match of size and form between pupae and adults, known larval synapomorphies in Staphylinidae and background knowledge of the Chilean fauna; they are described and illustrated. Structural and chaetotaxic characters of these and larvae of 25 other Omaliine Group genera were subjected to phylogenetic analysis. The position of Glypholomatinae within the Omaliine Group remains uncertain, although it is weakly supported as forming a clade with Omaliinae and Microsilphinae. Monophyly of the subfamily Proteininae and its components and of the tribe Coryphiini of Omaliinae are more strongly supported.