[1] Kyushu Univ, Grad Sch Bioresource & Bioenvironm Sci, Lab Adv Anim & Marine Bioresource, Fukuoka 8128581, Japan
[2] FANCL Corp, Cent Lab, Totsuka Ku, Yokohama, Kanagawa 2440806, Japan
[3] Virginia Polytech Inst & State Univ, Dept Anim & Poultry Sci, Blacksburg, VA 24061 USA