[1] Univ Penn, Sch Med, Ctr Neurodegenerat Dis Res, Philadelphia, PA 19104 USA
[2] Univ Penn, Sch Med, Dept Pathol & Lab Med, Philadelphia, PA 19104 USA
[3] Allegheny Univ Hlth Sci, Dept Neurol, Philadelphia, PA 19102 USA
[4] Univ Calif San Diego, Sch Med, Dept Neurosci, San Diego, CA 92103 USA