[1] Kyushu Univ, Fac Sci, Dept Biol, Higashi Ku, Fukuoka 8128581, Japan
[2] Gifu Univ, Sch Med, Dept Pediat, Gifu 5008076, Japan
[3] Himeji Inst Technol, Dept Life Sci, Kamigori, Hyogo 6781297, Japan
[4] Japan Sci & Technol Corp, CREST, Tokyo 1700013, Japan
[5] Johns Hopkins Univ, Sch Med, Kennedy Krieger Inst, Baltimore, MD 21205 USA
[6] Univ Amsterdam, Acad Med Ctr, Dept Pediat, NL-1100 DE Amsterdam, Netherlands