[1] Univ Oslo, Dept Biol, Div Zool, N-0316 Oslo, Norway
[2] London Sch Econ, Dept Stat, London WC2A 2AE, England
[3] Univ Hong Kong, Dept Stat & Actuarial Sci, Hong Kong, Hong Kong, Peoples R China
[4] Univ Warwick, Dept Stat, Coventry CV4 7AL, W Midlands, England