[1] Alfred Wegener Inst Polar & Marine Res, D-27570 Bremerhaven, Germany
[2] Univ Bremen, MARUM Ctr Marine Environm Sci, D-28359 Bremen, Germany
[3] Univ Bremen, Fac Geosci, D-28359 Bremen, Germany
[4] Max Planck Inst Marine Microbiol, Biogeochemistry Dept, D-28359 Bremen, Germany
[5] Univ Kiel IFM GEOMAR, Leibniz Inst Marine Sci, D-24148 Kiel, Germany
[6] Univ Tulsa, Dept Geosci, Tulsa, OK 74104 USA
[7] Adm Nacl Combustibles Alcoholes & Portland, Montevideo 11100, Uruguay