[1] Gifu Univ, Fac Agr, Gifu 5011193, Japan
[2] Kyoto Univ, Primate Res Inst, Inuyama, Aichi 4848506, Japan
[3] Natl Inst Infect Dis, Tsukuba Primate Ctr, Tsukuba, Ibaraki 3050843, Japan
[4] Sanwa Kagaku Kenkyusyo Co Ltd, Kumamoto 8693201, Japan
[5] Natl Inst Anim Hlth, Tsukuba, Ibaraki 3050856, Japan