[1] Johns Hopkins Univ, Dept Neurol, Sch Med, Baltimore, MD 21287 USA
[2] Johns Hopkins Univ, Dept Neurosci, Sch Med, Baltimore, MD 21287 USA
[3] Johns Hopkins Univ, Dept Physiol, Sch Med, Baltimore, MD 21287 USA
[4] Univ Texas, Ctr Dev Biol, Dallas, TX 75235 USA
[5] Yale Univ, Dept Neurol, New Haven, CT 06520 USA