[1] Univ Helsinki, Dept Forest Ecol, FIN-00014 Helsinki, Finland
[2] Swedish Univ Agr Sci, Dept Forest Ecol, S-90183 Umea, Sweden
[3] Mid Sweden Univ, Dept Chem & Proc Technol, S-85170 Sundsvall, Sweden
[4] Swedish Univ Agr Sci, Dept Forest Soil, S-75007 Uppsala, Sweden