[1] Tokyo Univ Fisheries, Dept Aquat Biosci, Minato Ku, Tokyo 1088477, Japan
[2] Kinki Univ, Fisheries Lab, Wakayama 6492211, Japan
[3] Mie Univ, Fac Bioresources, Tsu, Mie 5148507, Japan
[4] Keio Univ, Sch Med, Dept Mol Biol, Shinjuku Ku, Tokyo 1608582, Japan