[1] Univ British Columbia, Fac Agr Sci, Biometeorol & Soil Phys Grp, Vancouver, BC V6T 1Z4, Canada
[2] Univ Minnesota, Dept Soil Water & Climate, St Paul, MN USA
[3] Univ Guelph, Dept Land Resource Sci, Guelph, ON N1G 2W1, Canada
[4] British Columbia Minist Forests, Res Branch, Victoria, BC, Canada
[5] Univ Victoria, Dept Forest Biol, Victoria, BC, Canada