[1] Michigan State Univ, Ctr Microbial Ecol, E Lansing, MI 48824 USA
[2] Michigan State Univ, Dept Microbiol & Mol Genet, E Lansing, MI 48824 USA
[3] Michigan State Univ, Dept Civil & Environm Engn, E Lansing, MI 48824 USA
[4] Michigan State Univ, Dept Crop & Soil Sci, E Lansing, MI 48824 USA