[1] Univ Liverpool, Dept Pharmacol & Therapeut, Liverpool L69 3GE, Merseyside, England
[2] Univ Liverpool, Dept Dermatol, Liverpool L69 3GE, Merseyside, England
[3] Univ Bern, Inselspital, Clin Rheumatol & Clin Immunol Allergol, Bern, Switzerland
[4] Univ Liverpool, Dept Neurol Sci, Liverpool L69 3BX, Merseyside, England