[1] Univ Penn, Sch Med, Dept Med, Philadelphia, PA 19104 USA
[2] Univ Penn, Sch Med, Dept Genet, Philadelphia, PA 19104 USA
[3] Thomas Jefferson Univ, Dept Microbiol & Immunol Genet, Philadelphia, PA 19107 USA
[4] Thomas Jefferson Univ, Program Mol Biol, Philadelphia, PA 19107 USA