[1] Univ London, Queen Mary, William Harvey Res Inst, Dept Biochem Pharmacol, London EC1M 6BQ, England
[2] Univ Naples Federico II, Dipartimento Farmacol Sperimentale Federico II, Naples, Italy
[3] Univ Oxford, Dept Human Anat & Genet, Oxford OX1 3QX, England
[4] Imperial Coll Sch Med, Div Neurosci & Psychol Med, London W12 0NN, England