[1] Washington Univ, Sch Med, Ctr Immunol, St Louis, MO 63110 USA
[2] Washington Univ, Sch Med, Dept Pathol & Immunol, St Louis, MO 63110 USA
[3] Washington Univ, Sch Med, Howard Hughes Med Inst, St Louis, MO 63110 USA
[4] Washington Univ, Sch Med, Dept Med, Div Rheumatol, St Louis, MO 63110 USA