[1] Univ Calif Santa Cruz, Inst Marine Sci, Santa Cruz, CA 95064 USA
[2] Lamont Doherty Earth Observ, Palisades, NY 10964 USA
[3] Univ Hawaii, SOEST, Dept Oceanog, Honolulu, HI 96822 USA
[4] San Francisco State Univ, Romberg Tiburon Ctr, Tiburon, CA 94920 USA
[5] Univ So Calif, Wrigley Inst Marine Sci, Los Angeles, CA 90089 USA
[6] Univ So Calif, Dept Biol Sci, Los Angeles, CA 90089 USA
[7] Univ Calif Santa Cruz, Inst Marine Sci, Santa Cruz, CA 95064 USA