[1] Chinese Acad Sci, Inst Zool, Key Lab Zool Systemat & Evolut, Beijing 100101, Peoples R China
[2] Chinese Acad Sci, Inst Geog Sci & Nat Resources Res, Key Lab Ecosyst Network Observat & Modeling, Beijing 100101, Peoples R China
[3] Chinese Acad Sci, Inst Zool, State Key Lab Integrated Pest Management Pest Ins, Beijing 100101, Peoples R China
[4] Chinese Acad Sci, Grad Univ, Beijing 100101, Peoples R China