[1] Univ Saskatchewan, Sch Publ Hlth, Vaccinol & Immunotherapeut Program, 107 Wiggins Rd, Saskatoon, SK S7N 5E5, Canada
[2] Vaccine & Infect Dis Org, Int Vaccine Ctr, Saskatoon, SK S7N 5E3, Canada
[3] Univ Saskatchewan, Western Coll Vet Med, Dept Vet Microbiol, Saskatoon, SK S7N 5B4, Canada
[4] Univ Alberta, Edmonton, AB T6G 2J9, Canada