[1] Univ Kiel, Inst Nat Resource Conservat, D-24098 Kiel, Germany
[2] CSIRO Ecosyst Sci, Glen Osmond, SA 5064, Australia
[3] Bulgarian Acad Sci, Natl Inst Geophys Geodesy & Geog, BU-1113 Sofia, Bulgaria
[4] Wageningen Univ, Environm Syst Anal Grp, NL-6700 AA Wageningen, Netherlands
[5] PBL Netherlands Environm Assessment Agcy, NL-3720 AH Bilthoven, Netherlands