[1] Sainsbury Lab, Norwich NR4 7UH, Norfolk, England
[2] Biodivers & Climate Res Ctr BiK F, D-60325 Frankfurt, Germany
[3] Goethe Univ Frankfurt, Inst Ecol Evolut & Divers, Dept Biol Sci, D-60323 Frankfurt, Germany
[4] Massachusetts Inst Technol & Harvard, Broad Inst, Cambridge, MA 02141 USA
[5] Univ Delaware, Newark, DE 19716 USA