[1] Univ Calif Irvine, Dept Anat & Neurobiol, Irvine, CA 92697 USA
[2] Univ Calif Irvine, Dept Neurobiol & Behav, Irvine, CA USA
[3] Univ Calif Irvine, Dept Psychiat & Human Behav, Irvine, CA 92717 USA
[4] Univ So Calif, Neurosci Program, Los Angeles, CA 90089 USA