[1] Nagoya Univ, Ctr Gene Res, Chikusa Ku, Nagoya, Aichi 4648602, Japan
[2] Nagoya Univ, Biooriented Technol Res Adv Inst, Chikusa Ku, Nagoya, Aichi 4648602, Japan
[3] Nagoya Univ, Grad Sch Sci, Div Biol Sci, Chikusa Ku, Nagoya, Aichi 4648602, Japan
[4] Aichi Sci & Technol Fdn, Naka Ku, Nagoya, Aichi 4600002, Japan